In a display substrate and a vertical alignment display panel having the same, the display panel includes an array substrate and an opposite substrate. The color-forming layer includes a leuco dye that can be developed upon exposure to energy to produce a developed color. 0%, by weight, of polyelectolyte complexes having a compatible charge-balancing polymeric anion with at least 3 anionic groups and a total net charge of at least 4 negative charges.